Reflex Arc
In
biology, a type of action consisting of comparatively simple segments of
behaviour that usually occur as direct and immediate responses to particular stimuli
uniquely correlated with them. Reflex actions have a widespread occurrence
among complex animals.
Many
reflexes of placental mammals appear to be innate. They are transmitted by
heredity and are the common property of the species, and often of the genus.
They include not only such simple acts as mastication, swallowing, the blink
reflex, the knee jerk, and the scratch reflex, but also stepping, standing, the
cat's righting reflex, basic sexual acts, etc. Built up into complex patterns
of many coordinated muscular actions, reflexes form the basis of much instinctive
behaviour in animals.
Humans
also exhibit a variety of innate reflexes, which are variously concerned with
adjusting the musculature for optimum performance of the body's distance
receptors (the eye and the ear), with orienting parts of the body in spatial
relation to the head, and with managing the complicated acts involved in
ingesting food. Among the innate reflexes concerning just the eyes, for
example, are:
(1) paired
shifting of the eyeballs, often combined with turning of the head, to perceive
some interesting object in the field of vision;
(2) contraction
of the intraocular muscles to adjust the focus of the retina for the viewing of
near or far objects;
(3) constriction
of the pupil of the eye to reduce excessive illumination of the retina;
(4) blinking
due to intense light or touching of the cornea.
In
its simplest and most elementary form, a reflex is now viewed as a function of
an idealized mechanism called a reflex arc.
The primary components of the reflex arc have been identified as the
sensory-nerve cell (or receptor) that receives the stimulation, in turn
connecting to another nerve cell that activates the muscle cell (or effector),
which thus performs the reflex action. In most cases, however, the basic
physiological mechanism is more complicated than this simple arc theory would
suggest. Additional nerve cells capable of communicating with other parts of
the body (beyond the receptor and effector) are invariably present in reflex
circuits. As a result of the integrative action of the nervous system in higher
animals, the behaviour of such organisms is more than the simple sum of their
reflexes; it is a unitary whole that exhibits coordination between many
individual reflexes and is characterized not by inherited, stereotyped
responses but by flexibility and adaptability to circumstances.
